Until the Late Pleistocene, two further subspecies ranged across Europe and West Asia. These are U. t. mediterraneus from Western Europe and the Caucasus and U. t. permjak from Eastern Europe, particularly the Ural Mountains.[13]
Until the Late Pleistocene, two further subspecies ranged across Europe and West Asia. These are U. t. mediterraneus from Western Europe and the Caucasus and U. t. permjak from Eastern Europe, particularly the Ural Mountains.[13]