A 2016 study of Greenland shark eye tissue, published in the journal Science, estimated that these sharks can have a maximum life span of at least 272 years. The biggest shark in that study was estimated to be about 392 years old, and the researchers suggested that the sharks could have been up to 512 years old, Live Science previously reported. The age estimates came with a degree of uncertainty, but even the lowest estimate of 272 years still makes these sharks the longest-living vertebrates on Earth, Julius Nielsen, a researcher at Greenland Institute of Natural Resources, previously told Live Science.